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error 5.5.1 unable to send reply to email

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first occured today....i think that i changed a setting in the server info page but can't seem to recover/remember what it was. but now when i try to reply to a received email , i get the error message. please help!

first occured today....i think that i changed a setting in the server info page but can't seem to recover/remember what it was. but now when i try to reply to a received email , i get the error message. please help!